Marine microbes (phytoplankton) play an important role in the global cycling of elements including climatically significant carbon, sulfur, and nitrogen. Photosynthetic microbes in the surface ocean fix carbon and other elements into organic molecules, fueling food webs that sustain fisheries and most other life in the ocean. Sinking organic matter is remineralized and respired in the dark, sub-surface ocean maintaining a store of carbon about three times the size of the atmospheric inventory of CO2.